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Marx's theory of human nature occupies an important place in his critique of capitalism, his conception of communism, and his 'materialist conception of history'. Marx, however, does not refer to "human nature" as such, but to Gattungswesen, which is generally translated as 'species-being' or 'species-essence'. What Marx meant by this is that humans are capable of making or shaping their own nature to some extent. According to a note from the young Marx in the Manuscripts of 1844, the term is derived from Ludwig Feuerbach’s philosophy, in which it refers both to the nature of each human and of humanity as a whole . However, in the sixth Thesis on Feuerbach (1845), Marx criticizes the traditional conception of "human nature" as "species" which incarnates itself in each individual, on behalf of a conception of human nature as formed by the totality of "social relations". Thus, the whole of human nature is not understood, as in classical idealist philosophy, as permanent and universal: the species-being is always determined in a specific social and historical formation, with some aspects being biological.
Norman Geras claimed in Marx's theory of human nature (1983) that although many Marxists denied that there was a "human nature" to be found in Marx's words , there is in fact a Marxist conception of human nature which remains, to some degree, constant throughout history and across social boundaries. The sixth of the Theses on Feuerbach provided the basics for this interpretation of Marx according to which there was no eternal human nature to be found in his works. It states:
Feuerbach resolves the essence of religion into the essence of man [menschliche Wesen = ‘human nature’]. But the essence of man is no abstraction inherent in each single individual. In reality, it is the ensemble of the social relations. Feuerbach, who does not enter upon a criticism of this real essence is hence obliged:
- 1. To abstract from the historical process and to define the religious sentiment regarded by itself, and to presuppose an abstract — isolated - human individual.
- 2. The essence therefore can by him only be regarded as ‘species’, as an inner ‘dumb’ generality which unites many individuals only in a natural way. 
Thus, Marx appears to say that human nature is no more than what is made by the 'social relations'. Norman Geras's Marx's Theory of Human Nature, however, offers an extremely detailed argument against this position . In outline, Geras shows that, while the social relations are held to 'determine' the nature of people, they are not the only such determinant. In fact, Marx makes statements where he specifically refers to a human nature which is more than what is conditioned by the circumstances of one's life. In Capital, in a footnote critiquing utilitarianism, he says that utilitarians must reckon with 'human nature in general, and then with human nature as modified in each historical epoch' . Marx is arguing against an abstract conception of human nature, offering instead an account rooted in sensuous life. While he is quite explicit that '[a]s individuals express their life, so they are. Hence what individuals are depends on the material conditions of their production' , he also believes that human nature will condition (against the background of the productive forces and relations of production) the way in which individuals express their life. History involves 'a continuous transformation of human nature' , though this does not mean that every aspect of human nature is wholly variable; what is transformed need not be wholly transformed. Marx did criticise the tendency to 'transform into eternal laws of nature and of reason, the social forms springing from your present mode of production and form of property' , a process sometimes called "reification". For this reason, he would likely have wanted to criticise certain aspects of some accounts of human nature. Some people believe, for example, that humans are naturally selfish - Kant  and Hobbes  , for example. (Both Hobbes and Kant thought that it was necessary to constrain our human nature in order to achieve a good society - Kant thought we should use rationality, Hobbes thought we should use the force of the state - Marx, as we shall see, thought that the good society was one which allows our human nature its full expression.) Most Marxists will argue that this view is an ideological illusion and the effect of commodity fetishism: the fact that people act selfishly is held to be a product of scarcity and capitalism, not an immutable human characteristic. For confirmation of this view, we can see how, in The Holy Family Marx argues that capitalists are not motivated by any essential viciousness, but by the drive toward the bare 'semblance of a human existence' . (Marx says 'semblance' because he believes that capitalists are as alienated from their human nature under capitalism as the proletariat, even though their basic needs are better met.)
Man is directly a natural being. As a natural being and as a living natural being he is on the one hand endowed with natural powers, vital powers – he is an active natural being. These forces exist in him as tendencies and abilities – as instincts. On the other hand, as a natural, corporeal, sensuous objective being he is a suffering, conditioned and limited creature, like animals and plants. That is to say, the objects of his instincts exist outside him, as objects independent of him; yet these objects are objects that he needs – essential objects, indispensable to the manifestation and confirmation of his essential powers. 
In the Grundrisse Marx says his nature is a 'totality of needs and drives, which exerts a force upon me' . In The German Ideology he uses the formulation: 'their needs, consequently their nature' . We can see then, that from Marx's early writing to his later work, he conceives of human nature as composed of 'tendencies', 'drives', 'essential powers', and 'instincts' to act in order to satisfy 'needs' for external objectives. For Marx then, an explanation of human nature is an explanation of the needs of humans, together with the assertion that they will act to fulfill those needs. (c.f. The German Ideology, chapter 3 .) Norman Geras gives a schedule of some of the needs which Marx says are characteristic of humans:
...for other human beings, for sexual relations, for food, water, clothing, shelter, rest and, more generally, for circumstances that are conducive to health rather than disease. There is another one ... the need of people for a breadth and diversity of pursuit and hence of personal development, as Marx himself expresses these, 'all-round activity', 'all-round development of individuals', 'free development of individuals', 'the means of cultivating [one's] gifts in all directions', and so on. 
Marx says 'It is true that eating, drinking, and procreating, etc., are ... genuine human functions. However, when abstracted from other aspects of human activity, and turned into final and exclusive ends, they are animal.'  
In several passages throughout his work, Marx shows how he believes humans to be essentially different from other animals. 'Men can be distinguished from animals by consciousness, by religion or anything else you like. They themselves begin to distinguish themselves from animals as soon as they begin to produce their means of subsistence, a step which is conditioned by their physical organisation.'  In this passage from The German Ideology, Marx alludes to one difference: that humans produce their physical environments. But do not a few other animals also produce aspects of their environment as well? The previous year, Marx had already acknowledged:
In the same work, Marx writes:
Also in the segment on Estranged Labour:
More than twenty years later, in Capital, he came to muse on a similar subject:
From these passages we can observe something of Marx's beliefs about humans. That they characteristically produce their environments, and that they would do so, even were they not under the burden of 'physical need' - indeed, they will produce the 'whole of [their] nature', and may even create 'in accordance with the laws of beauty'. Perhaps most importantly, though, their creativity, their production is purposive and planned. Humans, then, make plans for their future activity, and attempt to exercise their production (even lives) according to them. Perhaps most importantly, and most cryptically, Marx says that humans make both their 'life activity' and 'species' the 'object' of their will. They relate to their life activity, and are not simply identical with it. Michel Foucault's definition of biopolitics as the moment when "man begins to take itself as a conscious object of elaboration" may be compared to Marx's definition hereby exposed.
To say that A is the object of some subject B, means that B (specified as an agent) acts upon A in some respect. Thus if 'the proletariat smashes the state' then 'the state' is the object of the proletariat (the subject), in respect of smashing. It is similar to saying that A is the objective of B, though A could be a whole sphere of concern and not a closely defined aim. In this context, what does it mean to say that humans make their 'species' and their 'lives' their 'object'? It's worth noting that Marx's use of the word 'object' can imply that these are things which humans produces, or makes, just as they might produce a material object. If this inference is correct, then those things that Marx says about human production above, also apply to the production of human life, by humans. And simultaneously, 'As individuals express their life, so they are. What they are, therefore, coincides with their production, both with what they produce and with how they produce. The nature of individuals thus depends on the material conditions determining their production.' 
To make one's life one's object is therefore to treat one's life as something that is under one's control. To raise in imagination plans for one's future and present, and to have a stake in being able to fulfill those plans. To be able to live a life of this character is to achieve 'self-activity' (actualisation), which Marx believes will only become possible after communism has replaced capitalism. 'Only at this stage does self-activity coincide with material life, which corresponds to the development of individuals into complete individuals and the casting-off of all natural limitations. The transformation of labour into self-activity corresponds to the transformation of the earlier limited intercourse into the intercourse of individuals as such' .
What is involved in making one's species one's object is more complicated (see Allen Wood 2004, pp16-21). In one sense, it emphasises the essentially social character of humans, and their need to live in a community of the species. In others, it seems to emphasise that we attempt to make our lives expressions of our species-essence; further that we have goals concerning what becomes of the species in general. The idea covers much of the same territory as 'making one's life one's object': it concerns self-consciousness, purposive activity, and so forth.
It is often said that Marx conceived of humans as homo faber, referring to Benjamin Franklin's definition of 'man as the tool-making animal' - that is, as 'man, the maker' , though he never used the term himself. Above, we indicated that one of Marx's central contentions about humans was that they were differentiated by the manner in which they produce and that thus, somehow, production was one of humans' essential activities. In this context, it is worth noting that Marx does not always address 'labour' or 'work' in such glowing terms. He says that communism 'does away with labour' . Furthermore, 'If it is desired to strike a mortal blow at private property, one must attack it not only as a material state of affairs, but also as activity, as labour. It is one of the greatest misapprehensions to speak of free, human, social labour, of labour without private property. “Labour” by its very nature is unfree, unhuman, unsocial activity, determined by private property and creating private property.'  Under Capitalism '[t]he capitalist functions only as capital personified, capital as a person, just as the worker only functions as the personification of labour, which belongs to him as torment, as exertion' .
It is generally held that Marx's view was that productive activity is an essential human activity, and can be rewarding when pursued freely. Marx's use of the words 'work' and 'labour' in the section above may be unequivocally negative; but this was not always the case, and is most strongly found in his early writing. However, Marx was always clear that under capitalism, labour was something inhuman, and dehumanising. 'labour is external to the worker – i.e., does not belong to his essential being; that he, therefore, does not confirm himself in his work, but denies himself, feels miserable and not happy, does not develop free mental and physical energy, but mortifies his flesh and ruins his mind' . While under communism, 'In the individual expression of my life I would have directly created your expression of your life, and therefore in my individual activity I would have directly confirmed and realised my true nature, my human nature, my communal nature' .
Marx's theory of history attempts to describe the way in which humans change their environments and (in dialectical relation) their environments change them as well. That is:
Further Marx set out his 'materialist conception of history' in opposition to 'idealist' conceptions of history; that of Hegel, for instance. 'The first premise of all human history is, of course, the existence of living human individuals. Thus the first fact to be established is the physical organisation of these individuals and their consequent relation to the rest of nature.'  Thus 'History does nothing, it “possesses no immense wealth”, it “wages no battles”. It is man, real, living man who does all that, who possesses and fights; “history” is not, as it were, a person apart, using man as a means to achieve its own aims; history is nothing but the activity of man pursuing his aims' . So we can see that, even before we begin to consider the precise character of human nature, 'real, living' humans, 'the activity of man pursuing his aims' is the very building block of Marx's theory of history. Humans act upon the world, changing it and themselves; and in doing so they 'make history' . But even beyond this, human nature plays two key roles. In the first place, it is part of the explanation for the growth of the productive forces, which Marx conceives of as the driving force of history. Secondly, the particular needs and drives of humans explain the class antagonism which is generated under capitalism.
It has been held by several writers that it is Marx's conception of human nature which explains the 'primacy thesis' (Cohen, 1978) concerning the expansion of the productive forces, which according to Marx, is itself the fundamental driving force of history. If true, this would make his account of human nature perhaps the most fundamental aspect of his work. Geras writes, (1983, p107-108, italics in original) 'historical materialism itself, this whole distinctive approach to society that originates with Marx, rests squarely upon the idea of a human nature. It highlights that specific nexus of universal needs and capacities which explains the human productive process and man's organized transformation of the material environment; which process and transformation it treats in turn as the basis both of the social order and of historical change.' G.A. Cohen (1988, p84): 'The tendency's autonomy is just its independence of social structure, its rootedness in fundamental material facts of human nature and the human situation.' Allen Wood (2004, p75): 'Historical progress consists fundamentally in the growth of people's abilities to shape and control the world about them. This is the most basic way in which they develop and express their human essence' (see also, the quotation from Allen Wood above).
In his article Reconsidering Historical Materialism, however, Cohen gives an argument to the effect that human nature cannot be the premise on which the plausibility of the expansion of the productive forces is grounded.
The implication of this is that hence 'one might ... imagine two kinds of creature, one whose essence it was to create and the other not, undergoing similarly toilsome histories because of similarly adverse circumstances. In one case, but not the other, the toil would be a self-alienating exercise of essential powers' (p170). Hence, 'historical materialism and Marxist philosophical anthropology are independent of, though also consistent with, each other' (p174, see especially sections 10 and 11). The problem is this: it seems as though the motivation most people have for the work they do isn't the exercise of their creative capacity; on the contrary, labour is alienated by definition in the capitalist system based on salary, and people only do it because they have to. They go to work not to express their human nature but to find theirs means of subsistence So in that case, why do the productive forces grow - does human nature have anything to do with it? The answer to this question is a difficult one, and a closer consideration of the arguments in the literature is necessary for a full answer than can be given in this article. However, it is worth bearing in mind that Cohen had previously been committed to the strict view that human nature (and other 'asocial premises') were sufficient for the development of the productive forces - it could be that they are only one necessary constituent. It is also worth considering that by 1988 (see quotation above), he appears to consider that the problem is resolved.
Some needs are far more important than others. In The German Ideology Marx writes that 'life involves before everything else eating and drinking, a habitation, clothing and many other things'. All those other aspects of human nature which he discusses (such as 'self-activity') are therefore subordinate to the priority given to these. Marx makes explicit his view that humans develop new needs to replace old: 'the satisfaction of the first need (the action of satisfying, and the instrument of satisfaction which has been acquired) leads to new needs' .
Geras says of Marx's work that: 'Whatever else it is, theory and socio-historical explanation, and scientific as it may be, that work is a moral indictment resting on the conception of essential human needs, an ethical standpoint, in other words, in which a view of human nature is involved' (1983, p83-84).
Alienation, for Marx, is the estrangement of humans from aspects of their human nature. Since - as we have seen - human nature consists in a particular set of vital drives and tendencies, whose exercise constitutes flourishing, alienation is a condition wherein these drives and tendencies are stunted. For essential powers, alienation substitutes disempowerment; for making one's own life one's object, one's life becoming an object of capital. Marx believes that alienation will be a feature of all society before communism. The opposite of, alienation is 'actualisation' or 'self-activity' - the activity of the self, controlled by and for the self.
One important criticism of Marx's 'philosophical anthropology' (i.e. his conception of humans) is offered by Gerald Cohen, the leader of "Analytical Marxism", in Reconsidering Historical Materialism (in ed. Callinicos, 1989). Cohen claims: 'Marxist philosophical anthropology is one sided. Its conception of human nature and human good overlooks the need for self identity than which nothing is more essentially human.' (p173, see especially sections 6 and 7). The consequence of this is held to be that 'Marx and his followers have underestimated the importance of phenomena, such as religion and nationalism, which satisfy the need for self identity. (Section 8.)' (p173). Cohen describes what he sees as the origins of Marx's alleged neglect: 'In his anti-Hegelian, Feuerbachian affirmation of the radical objectivity of matter, Marx focused on the relationship of the subject to an object which is in no way subject, and, as time went on, he came to neglect the subject's relationship to itself, and that aspect of the subject's relationship to others which is a mediated (that is, indirect), form of relationship to itself' (p155).
Consequently, Cohen believes, 'A person does need to develop and ENJOY his POWERS, he needs to gain and gain until he is the best and no one else can overpower him, people are BAD" He must, as Hegel saw, find something outside himself which he did not create, and to which something inside himself corresponds, because of the social process that created him' (p156). Cohen believes that people are driven, typically, not to create identity, but to preserve that which they have in virtue, for example, of 'nationality, or race, or religion, or some slice or amalgam thereof' (p156-159). Cohen does not claim that 'Marx denied that there is a need for self definition, but [instead claims that] he failed to give the truth due emphasis' (p155). Nor does Cohen say that the sort of self understanding that can be found through religion etc. is accurate (p158). Of nationalism, he says 'identifications [can] take benign, harmless, and catastrophically malignant forms' (p157) and does not believe 'that the state is a good medium for the embodiment of nationality' (p164).
All the quotations from Marx in this article have used the translation employed by the Marxists Internet Archive. This means that you can follow the external reference links, and then search on that page using your browser's search function for some part of the text of the quotation in order to ascertain its context.
The two texts in which Marx most directly discusses human nature are the Comments on James Mill and the piece on Estranged Labour in the Economic and Philosophical Manuscripts of 1844 (published in 1932). Both of these pieces date from 1844, and as such were written by the young Marx; some analysts (Louis Althusser, etc.) assert that work from this period differs markedly in its ideas from the later work.
In certain aspects, the views of many earlier writers on this topic are generally believed to have been superseded. Nevertheless, here is a selection of the best writing prior to 1978. Much of it addresses human nature through the strongly related concept of alienation: